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Reference: H.H. Tan & P. Ng, 2006. Ichthyol. Explor. Freshwaters 17 (2)
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Tan, H.H. & Ng, P. 2006. Six new species of fighting fish (Teleostei: Osphronemidae: Betta) from Borneo. Ichthyol. Explor. Freshwaters 17 (2): 97-114.

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