[organisms] see Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
   Domain Bacteria Woese et al. 1990
     Phylum Nitrospira Garrity & Holt, 2001 see Phylum Nitrospirae Garrity and Holt 2001
       Class "Nitrospira" 2001
         Order "Nitrospirales" 2001
           Family "Nitrospiraceae" 2001
             Genus Leptospirillum (ex Markosyan 1972) Hippe 2000
               1 Leptospirillum ferrooxidans (ex Markosyan 1972) Hippe 2000
               2 Leptospirillum ferriphilum Coram and Rawlings 2002
               3 Leptospirillum thermoferrooxidans Hippe 2000