Natura - nature
   actualia - actual entities
     Mundus Plinius - physical world
       naturalia - natural bodies
         Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
           Domain Bacteria Woese et al. 2024
             Kingdom Pseudomonadati (Gibbons and Murray 1978) Oren and Göker 2024
               Phylum Pseudomonadotaᵀ Garrity et al. 2021
                 Class Hydrogenophilia Boden et al. 2022
                   Order Hydrogenophilalesᵀ Garrity et al. 2006 [monotypic]
                     Family Hydrogenophilaceaeᵀ Garrity et al. 2006
                       Genus Tepidiphilus Manaia et al. 2003 [monotypic]
                         Tepidiphilus margaritiferᵀ Manaia et al. 2003