Natura - nature
   actualia - actual entities
     Mundus Plinius - physical world
       naturalia - natural bodies
         Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
           Clade neomura Cavalier-Smith, 1987
             Domain Eukaryota Chatton, 1925 - eukaryotes
               Clade discaria Cavalier-Smith, 2021
                 Clade dorsates Cavalier-Smith, 2021
                   Clade podiates Cavalier-Smith, 2012
                     Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
                       Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
                         Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
                           Superkingdom Holomycota Liu, Steenkamp, Brinkmann, Forget, Philippe & Lang, 2009
                             Kingdom Fungi R.T.Moore (1980) - fungi
                               Subkingdom Dikarya Hibbett, T.Y. James & Vilgalys (2007)
                                 Phylum Basidiomycota R.T.Moore (1980) - basidiomycetes
                                   Subphylum Pucciniomycotina R. Bauer, Begerow, J.P. Samp., M. Weiss & Oberw. (2006)
                                     Class Pucciniomycetesᵀ R. Bauer, Begerow, J.P. Samp., M. Weiss & Oberw. (2006)
                                       Order Puccinialesᵀ Clem. & Shear (1931)
                                         Family Pucciniastraceae Gäum. ex Leppik (1972) [paraphyletic]
                                           01 Genus Hyalopsora Magnus (1902)
                                           02 Genus Melampsorella J.Schröt. (1874)
                                           03 Genus Melampsoridium Kleb. (1899)
                                           04 Genus Milesina Magnus (1909)
                                           05 Genus Naohidemyces S. Sato, Katsuya & Y. Hirats. (1993)
                                           06 Genus Pucciniastrumᵀ G.H. Otth (1861)
                                           07 Genus Thekopsora Magnus (1875)
                                           08 Genus Uredinopsis Magnus (1893)
                                           09 Anamorph genus Milesia F.B. White (1878)
                                           10 Anamorph genus Peridiopsora Kamat & Sathe (1969)