Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Order Coleoptera Linnaeus, 1758 - beetles
Suborder Adephaga Schellenberg, 1806
Family Carabidae Latreille, 1802 - ground beetles
Subfamily Carabinaeᵀ Latreille, 1802
Genus Carabusᵀ Linnaeus, 1758
Subgenus Carabus (Archicarabus) {Seidlitz, 1887}
Subgenus Carabus (Callistocarabus) {Reitter, 1896}
Subgenus Carabus (Carabus)ᵀ Linnaeus, 1758
Carabus angulicollis
Carabus limbatus Say, 1825
Carabus olympiae Sella, 1855
Carabus scabrosus G.A.Olivier, 1790
Carabus serratus Say, 1823
Carabus taedatus Fabricius, 1787
Carabus vinctus (Weber, 1801)
Subgenus Carabus (Chaetocarabus) {Thomson, 1875}
Subgenus Carabus (Chrysocarabus) {Thomson, 1875}
Subgenus Carabus (Eucarabus) {Géhin, 1876}
Subgenus Carabus (Hemicarabus) {Géhin, 1885}
Subgenus Carabus (Hygrocarabus) {Thomson, 1875}
Subgenus Carabus (Megodontus) Solier, 1848
Subgenus Carabus (Mesocarabus) {Thomson, 1875}
Subgenus Carabus (Oreocarabus) {Géhin, 1885}
Subgenus Carabus (Orinocarabus) {Kraatz, 1878}
Subgenus Carabus (Pachystus) {Motschulsky, 1865}
Subgenus Carabus (Platycarabus) {Morawitz, 1886}
Subgenus Carabus (Procrustes) Bonelli, 1809
Subgenus Carabus (Tomocarabus) Reitter, 1896
Subgenus Carabus (Trachycarabus) Géhin, 1885