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Taxon: Order Lepidoptera Linnaeus, 1758 - butterflies, moths (moth)
Classification by:

Natura - nature
Ref: C. Linnaeus, 1748: 210; Count: 14k;226p;874c;1605o;9906f;114,342g;258,955s;29,252ss;2100v;35,690;112e; ‡2o;20f;118g;518s;110ss;1v;7; †4p;75c;550o;3448f;17,150g;6300s;207ss;2v;392;14e
   actualia - actual entities
  Ref: A. Voltolini, 2006: 44 (note 19) [assumed position]; Count: 14k;226p;874c;1605o;9906f;114,342g;258,954s;29,252ss;2100v;35,690;112e; ‡2o;20f;118g;518s;110ss;1v;7; †4p;75c;550o;3448f;17,150g;6300s;207ss;2v;392;14e
     Mundus Plinius - physical world
    Ref: C. Linnaeus, 1758: 5 [indirect position]; 24 Apr 2014; Count: 14k;226p;874c;1605o;9906f;114,342g;258,954s;29,252ss;2100v;35,688;112e; ‡2o;20f;118g;518s;110ss;1v;7; †4p;75c;550o;3448f;17,150g;6300s;207ss;2v;392;14e
       naturalia - natural bodies
      Ref: C. Linnaeus, 1758: 6 [implicit position]; Count: 8k;226p;874c;1605o;9830f;114,332g;258,530s;29,252ss;2069v;9; ‡2o;20f;118g;518s;110ss;1v; †4p;75c;550o;3448f;17,150g;6300s;207ss;2v
         [crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
        Ref: J. Wiemann et al., 2020 [validation] [inferred position]; Count: 6k;226p;399c;1597o;9830f;114,289g;255,303s;29,250ss;2041v;9; ‡2o;20f;118g;518s;110ss;1v; †4p;75c;549o;3448f;17,141g;6260s;207ss;2v
           Clade "neomura" Cavalier-Smith 1987
          Ref: T. Cavalier-Smith, 2002b: 297 [inferred position]; Count: 6k;81p;328c;1441o;9332f;110,744g;238,979s;28,849ss;2023v;2; ‡2o;20f;118g;518s;110ss;1v; †4p;75c;549o;3447f;17,134g;6251s;207ss;2v
             Domain Eukaryota Chatton, 1925 - eukaryotes
            Ref: P.M. Kirk et al., 2008: 248 [indirect position]; 19 Sep 2015; Count: 6k;76p;318c;1422o;9300f;110,617g;238,533s;28,849ss;2023v;2; ‡2o;20f;118g;518s;110ss;1v; †4p;75c;549o;3447f;17,134g;6251s;207ss;2v
               Clade neokaryotes Cavalier-Smith 1993 - mitochondrial eukaryotes
              Ref: T. Cavalier-Smith, 2013(2012): 116 [inferred position]; 18 Sep 2015; Count: 5k;75p;314c;1414o;9294f;110,399g;238,360s;28,842ss;2023v;2; ‡2o;20f;118g;518s;110ss;1v; †4p;75c;549o;3444f;17,040g;6165s;201ss;2v
                 Clade neozoa Cavalier-Smith 1993
                Ref: T. Cavalier-Smith, 2013(2012): fig. 2; Count: 5k;74p;308c;1406o;9283f;110,362g;238,305s;28,842ss;2023v;2; ‡2o;20f;118g;518s;110ss;1v; †4p;75c;549o;3444f;17,040g;6165s;201ss;2v
                   Clade scotokaryotes Cavalier-Smith 2000
                  Ref: T. Cavalier-Smith et al., 2015: 357 [emended diagnosis] [indirect position]; 18 Sep 2015; Count: 3k;56p;192c;872o;7292f;88,644g;217,163s;28,335ss;511v;1; ‡2o;20f;117g;514s;110ss;1v; †4p;70c;491o;3021f;13,264g;5068s;107ss;2v
                     Clade podiates Cavalier-Smith 2012
                    Ref: T. Cavalier-Smith et al., 2015: 357 [new position]; Count: [p] 3k;54p;186c;858o;7256f;88,529g;217,004s;28,335ss;511v;1; ‡2o;20f;117g;514s;110ss;1v; †4p;70c;491o;3021f;13,264g;5068s;107ss;2v
                       [crown] Clade Amorphea Adl et al. 2012
                      Ref: S.M. Adl et al., 2012: table 1 (p. 430) [new taxon] [indirect position]; 19 Sep 2015; Count: [p] 3k;54p;183c;855o;7251f;88,520g;216,991s;28,335ss;511v;1; ‡2o;20f;117g;514s;110ss;1v; †4p;70c;491o;3021f;13,264g;5068s;107ss;2v
                         [crown] Clade Opisthokonta Cavalier-Smith 1987
                        Ref: E.J. van Nieukerken, 2010: 98 [indirect position]; Count: [p] 3k;52p;174c;824o;7176f;88,275g;216,530s;28,333ss;507v;1; ‡2o;20f;117g;514s;110ss;1v; †4p;70c;491o;3021f;13,264g;5067s;107ss;2v
                           Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
                          Ref: L. Tedersoo et al., 2018: 154 [implicit position]; Count: [p] 1k;33p;96c;582o;6241f;78,960g;214,166s;28,325ss;483v;1; ‡2o;20f;117g;514s;110ss;1v; †4p;69c;490o;3020f;12,778g;5058s;107ss;2v
                             Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith 2008
                            Ref: T. Cavalier-Smith, 2013(2012): 156 [inferred position]; 19 Jan 2016; Count: [p] 1k;33p;95c;580o;6234f;78,940g;214,144s;28,325ss;483v;1; ‡2o;20f;117g;514s;110ss;1v; †4p;69c;490o;3020f;12,778g;5058s;107ss;2v
                               Kingdom Animalia Linnaeus, 1758 - animals
                              Ref: T. Cavalier-Smith, 2013(2012): 156 [implicit position]; Count: [p] 33p;92c;576o;6228f;78,884g;214,076s;28,325ss;482v;1; ‡2o;20f;117g;514s;110ss;1v; †4p;69c;490o;3020f;12,778g;5058s;107ss;2v
                                 Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
                                Ref: G. Giribet & G.D. Edgecombe, 2019: 168 (fig. 1a) [inferred position]; Giribet & Edgecombe, 2019: fig. 1a; Count: 31p;86c;541o;6063f;78,121g;213,526s;28,306ss;471v;1; ‡2o;20f;117g;514s;110ss;1v; †2p;63c;461o;2723f;11,278g;4804s;107ss;2v
                                   Subkingdom Bilateria Hatschek, 1888 - bilaterians
                                  Ref: M.A. Ruggiero et al., 2015b: table 2 (p. 36) [indirect position]; Giribet & Edgecombe, 2019: fig. 1a; Count: 29p;80c;518o;5826f;77,229g;212,789s;28,301ss;463v;1; ‡2o;20f;117g;514s;110ss;1v; †2p;58c;447o;2710f;11,154g;4708s;107ss;2v
                                     Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
                                    Ref: G. Giribet & G.D. Edgecombe, 2019: 168 (fig. 1b) [implicit position]; Giribet & Edgecombe, 2019: fig. 1b; Count: 28p;78c;516o;5802f;77,143g;212,767s;28,301ss;463v;1; ‡2o;20f;117g;514s;110ss;1v; †1p;55c;447o;2706f;11,134g;4686s;107ss;2v
                                       Infrakingdom Protostomia Grobben, 1908
                                      Ref: M.A. Ruggiero et al., 2015b: table 2 (p. 36) [indirect position]; Giribet & Edgecombe, 2019: fig. 1b; Count: 25p;60c;326o;4554f;66,649g;150,654s;15,594ss;155v; ‡1f;2g;101s;8ss; †27c;180o;1539f;3545g;1639s;32ss;1v
                                         Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
                                        Ref: M.A. Ruggiero et al., 2015b: table 2 (p. 36); Giribet & Edgecombe, 2019: fig. 1b; Count: [p] 8p;27c;169o;3281f;57,822g;134,943s;15,149ss;151v; ‡1f;2g;57s;4ss; †15c;91o;618f;2379g;1345s;20ss
                                           Panarthropoda Nielsen, 1995 - panarthropods
                                          Ref: E.J. van Nieukerken, 2010: 152 [implicit position]; Count: [p] 3p;19c;130o;2995f;57,082g;133,973s;15,143ss;151v; ‡1f;2g;57s;4ss; †14c;91o;617f;2340g;1340s;20ss
                                             Tactopoda Budd, 2001
                                            Ref: R.J. Howard et al., 2020: fig. S1[d] [implicit position]; Count: 2p;18c;129o;2993f;57,063g;133,957s;15,143ss;151v; ‡1f;2g;57s;4ss; †13c;87o;606f;2319g;1315s;20ss
                                               Phylum Arthropoda Siebold, 1848 - arthropods
                                              Ref: R.J. Howard et al., 2020: fig. S1[d]; Count: 15c;124o;2970f;56,968g;133,576s;15,127ss;148v; ‡1f;2g;57s;4ss; †13c;87o;605f;2318g;1313s;20ss
                                                 Euarthropoda
                                                Ref: J.C. Lamsdell, 2013: 14 [implicit position]; Count: 15c;124o;2970f;56,968g;133,576s;15,127ss;148v; ‡1f;2g;57s;4ss; †9c;74o;586f;2281g;1297s;20ss
                                                   Antennulata
                                                  Ref: C. Aria et al., 2020: 8 [implicit position]; Count: 12c;106o;2189f;46,788g;108,959s;14,357ss;146v; ‡1g;57s;4ss; †6c;61o;463f;1764g;1195s;16ss
                                                     [Subphylum] Mandibulata Snodgrass, 1938
                                                    Ref: C. Aria et al., 2020: 8 [inferred rank]; Count: 12c;106o;2189f;46,788g;108,959s;14,357ss;146v; ‡1g;57s;4ss; †3c;43o;288f;1132g;1067s;10ss
                                                       Infraphylum Pancrustacea Zrzavý & Štys, 1997
                                                      Ref: Schweitzer et al., 2020: 246 [Edgecombe, 2017: fig. 1] [indirect position]; Count: 8c;85o;2037f;46,085g;108,300s;14,326ss;105v; ‡1g;57s;4ss; †1c;34o;280f;1103g;1029s;10ss
                                                         Altocrustacea
                                                        Ref: J.C. Regier et al., 2010: fig. 1 [implicit position]; Count: 7c;77o;1968f;45,319g;107,804s;14,320ss;102v; ‡1g;55s;4ss; †1c;30o;263f;998g;955s;10ss
                                                           Miracrustacea
                                                          Ref: J.C. Regier et al., 2010: fig. 1; Count: 5c;38o;1093f;38,290g;101,265s;14,163ss;102v; ‡1g;46s;4ss; †16o;254f;891g;928s;10ss
                                                             Superclass Hexapoda Latreille, 1825
                                                            Ref: E.E. Ruppert et al., 2004: 723 [indirect position]; Count: 3c;36o;1089f;38,277g;101,255s;14,163ss;102v; ‡1g;46s;4ss; †15o;253f;887g;925s;10ss
                                                               Cercophora
                                                              Ref: G. Giribet & G.D. Edgecombe, 2020: 235 [implicit position]; Count: 2c;31o;1051f;38,115g;100,339s;14,161ss;102v; ‡1g;46s;4ss; †15o;250f;885g;923s;10ss
                                                                 [Class] Insecta Linnaeus, 1758 - insects
                                                                Ref: G. Giribet & G.D. Edgecombe, 2020: 235 [inferred rank]; Count: 31o;1051f;38,115g;100,339s;14,161ss;102v; ‡1g;46s;4ss; †15o;250f;885g;923s;10ss
                                                                   Subclass Dicondylia Haeckel, 1866
                                                                  Ref: R.W. Poole et al., 1997: 16; Count: [oo] 30o;1049f;38,096g;100,310s;14,161ss;102v; ‡1g;46s;4ss; †15o;248f;881g;919s;10ss
                                                                     Infraclass Pterygota Lang, 1888
                                                                    Ref: R.W. Poole et al., 1997: 16; Count: [oo] 29o;1044f;38,077g;100,288s;14,161ss;102v; ‡1g;46s;4ss; †15o;248f;878g;916s;10ss
                                                                       Metapterygota
                                                                      Ref: D. Grimaldi & M.S. Engel, 2005: table 4.1 [implicit position]; Count: 28o;1016f;37,783g;99,450s;14,140ss;102v; ‡1g;44s;4ss; †14o;227f;829g;846s;10ss
                                                                         Neoptera Martynov, 1923
                                                                        Ref: D. Grimaldi & M.S. Engel, 2005: table 4.1; Count: 27o;987f;37,152g;98,886s;14,110ss;102v; ‡1g;44s;4ss; †8o;171f;825g;843s;10ss
                                                                           Eumetabola
                                                                          Ref: D. Grimaldi & M.S. Engel, 2005: table 4.1; Count: 17o;863f;33,613g;91,060s;12,881ss;102v; ‡1g;43s;4ss; †4o;123f;529g;823s;10ss
                                                                             Holometabola
                                                                            Ref: D. Grimaldi & M.S. Engel, 2005: table 4.1; Count: [o] 13o;657f;30,450g;89,488s;12,829ss;101v; ‡1g;41s;4ss; †2o;112f;507g;813s;10ss
                                                                               Superorder Panorpida
                                                                              Ref: D. Grimaldi & M.S. Engel, 2005: table 4.1; Count: [o] 8o;370f;19,962g;71,439s;10,691ss;7v; ‡24s;4ss; †22f;162g;26s
                                                                                 Amphiesmenoptera
                                                                                Ref: D. Grimaldi & M.S. Engel, 2005: table 4.1; Count: [f] 2o;179f;17,124g;67,412s;10,626ss;1v; ‡21s;4ss; †17f;155g;14s
                                                                                   Order Lepidoptera Linnaeus, 1758 - butterflies, moths
                                                                                  Ref: D. Grimaldi & M.S. Engel, 2005: table 4.1; Count: [f] 133f;16,516g;67,344s;10,623ss;1v; ‡21s;4ss; †4f;55g;13s
                                                                                     01 Genus †Archiptilia Handlirsch, 1939, incertae sedis
                                                                                    Ref: E.J. van Nieukerken et al., 2011: 212 (footnote 4); Count: -
                                                                                     02 Genus †Electrocrania Kuznetzov, 1941, incertae sedis [monotypic]
                                                                                    Ref: E.J. van Nieukerken et al., 2011: 212 (footnote 4); Count: [†*] †1s
                                                                                     03 Genus †Epididontus Handlirsch, 1939, incertae sedis
                                                                                    Ref: E.J. van Nieukerken et al., 2011: 212 (footnote 4); Count: -
                                                                                     04 Genus †Palaeosabatinca Kozlov, 1988, incertae sedis
                                                                                    Ref: E.J. van Nieukerken et al., 2011: 212 (footnote 4); Count: -
                                                                                     05 Genus †Parataulius Handlirsch, 1939, incertae sedis
                                                                                    Ref: E.J. van Nieukerken et al., 2011: 212 (footnote 4); Count: -
                                                                                     06 Genus †Phylledestes Cockerell, 1907, incertae sedis [monotypic]
                                                                                    Ref: G. Beccaloni et al., 2005: Feb 2005; Count: [†*] †1s
                                                                                     07 Genus †Xena Martins-Neto, 1999, nom. illeg., incertae sedis
                                                                                    Ref: E.J. van Nieukerken et al., 2011: 212 (footnote 4); Count: -
                                                                                     08 Family †Archaeolepidae Whalley, 1985, incertae sedis [monotypic]
                                                                                    Ref: E.J. van Nieukerken et al., 2011: 212 [Grimaldi & Engel, 2005: table 13.1]; Count: [†*] †1g;1s
                                                                                     09 Family †Mesokristenseniidae Huang, Nel & Minet, 2010, incertae sedis [monotypic]
                                                                                    Ref: E.J. van Nieukerken et al., 2011: 212; Count: -
                                                                                     10.1.1 Genus †Auliepterix Kozlov, 1989, incertae sedis
                                                                                    Ref: D. Grimaldi & M.S. Engel, 2005: table 13.1; Count: †1s
                                                                                     10.1.2 Family †Eolepidopterigidae Rasnitsyn, 1983, incertae sedis [monotypic]
                                                                                    Ref: E.J. van Nieukerken et al., 2011: 212 [Grimaldi & Engel, 2005: table 13.1]; Count: [†*] †1g;1s
                                                                                     10.1.3 Genus †Karataunia Kozlov, 1989, incertae sedis
                                                                                    Ref: D. Grimaldi & M.S. Engel, 2005: table 13.1; Count: -
                                                                                     10.1.4 Genus †Palaeolepidopterix Kozlov, 1989, incertae sedis
                                                                                    Ref: D. Grimaldi & M.S. Engel, 2005: table 13.1; Count: -
                                                                                     10.1.5 Genus †Protolepis Kozlov, 1989, incertae sedis
                                                                                    Ref: D. Grimaldi & M.S. Engel, 2005: table 13.1; Count: -
                                                                                     10.2.1 Family †Undopterigidae Kozlov, 1988, incertae sedis [monotypic]
                                                                                    Ref: E.J. van Nieukerken et al., 2011: 212; Count: [†*] †1g;2s
                                                                                     10.2.2 Genus †Daiopterix Skalski, 1984, incertae sedis
                                                                                    Ref: D. Grimaldi & M.S. Engel, 2005: table 13.1; Count: †2s
                                                                                     10.2.3 Genus †Gracilepterix Martins-Neto & Vulcano, 1989, incertae sedis
                                                                                    Ref: D. Grimaldi & M.S. Engel, 2005: table 13.1; Count: †1s
                                                                                     10.3.1.1 Genus †Parasabatinca Whalley, 1978, incertae sedis
                                                                                    Ref: D. Grimaldi & M.S. Engel, 2005: table 13.1; Count: [†*] †2s
                                                                                     10.3.1.2 Suborder Zeugloptera Chapman, 1917 [monotypic]
                                                                                    Ref: E.J. van Nieukerken et al., 2011: 212; Count: [*] 1f;10g;119s;8ss
                                                                                     10.3.2.1 Suborder Aglossata Speidel, 1977 [monotypic]
                                                                                    Ref: E.J. van Nieukerken et al., 2011: 212; Count: [*] 1f;1g;2s
                                                                                     10.3.2.2.1 Suborder Heterobathmiina Kristensen & Nielsen, 1983 [monotypic]
                                                                                    Ref: E.J. van Nieukerken et al., 2011: 212; Count: [*] 1f;1g;2s
                                                                                     10.3.2.2.2 Suborder Glossata Fabricius, 1775
                                                                                    Ref: E.J. van Nieukerken et al., 2011: 212; Count: [f] 130f;16,504g;67,221s;10,615ss;1v; ‡21s;4ss; †38g;1s

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Last updated: 4 Aug 2022