Kingdom Animalia Linnaeus, 1758 - animals
   Phylum Arthropoda Siebold, 1848 - arthropods
     Class Insecta Linnaeus, 1758 - insects
       Order Lepidoptera Linnaeus, 1758 - butterflies, moths
         Family Nymphalidae Rafinesque, 1815 - brush-footed butterflies
           Genus Boloria Moore, 1900