Phylum Arthropoda Siebold, 1848 - arthropods
   Class Insecta Linnaeus, 1758 - insects
     Subclass Pterygota Lang, 1888 - winged insects see Infraclass Pterygota Lang, 1888
       Endopterygota Sharp, 1899
         Order Lepidoptera Linnaeus, 1758 - butterflies, moths
           Suborder Heteroneura Tillyard, 1918 see Infraorder Heteroneura Tillyard, 1918
             Superfamily Papilionoidea Latreille, 1802
               Family Satyridae Boisduval, 1833 see Subfamily Satyrinae Boisduval, 1833
                 Subfamily Satyrinae Boisduval, 1833
                   Tribe Maniolini see Subtribe Maniolina Grote, 1897
                     1 Genus Cercyonis Speyer ex Scudder, 1875
                     2 Genus Hyponephele Muschamp, 1915
                     3 Genus Maniola Schrank, 1801
                     3.0 Genus Epinephele Hübner, 1819 see Genus Maniola Schrank, 1801
                     4 Genus Pyronia Hübner, 1819