[organisms] see Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
   Domain "Archaea" Woese et al. 1990
     Phylum "Euryarchaeota" Garrity and Holt 2001
       Class "Methanomicrobia"
         Order Methanomicrobiales Balch and Wolfe 1981
           Family Methanomicrobiaceae Balch and Wolfe 1981
             Genus Methanoculleus Maestrojuán et al. 1990
               Methanoculleus bourgensis corrig. (Ollivier et al. 1986) Maestrojuán et al. 1990