Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Order Coleoptera Linnaeus, 1758 - beetles
Suborder Polyphaga Emery, 1886
Series Scarabaeiformia Crowson, 1960
Superfamily Scarabaeoideaᵀ Latreille, 1802
Family Scarabaeidaeᵀ Latreille, 1802 - scarab beetles, dung beetles, and june "bugs"
Subfamily Aphodiinae Leach, 1815
Genus Annegialia Howden, 1971
Genus Aphotaenius Cartwright, 1952
Genus Dialytellus Brown, 1929
Genus Dialytes Harold, 1869
Genus Euparixia Brown, 1927
Genus Geopsammodius Gordon & Pittino, 1992
Genus Leiopsammodius Rokovic, 1979
Genus Neopsammodius Rakovic, 1986
Genus Odontopsammodius Gordon & Pittino, 1992
Genus Platytomus Mulsant, 1842
Genus Pseudataenius Brown, 1927
Genus Termitaxis J. Krikken, 1970
Genus Tesarius Rokovic, 1981
Genus Trichiorhyssemus Clouet, 1901
Genus Xeropsamobeus Saylor, 1937
01 Tribe Aphodiiniᵀ Leach, 1815
02 Tribe Corythoderini Schmidt, 1910
03 Tribe Eupariini Schmidt, 1910, nom. protect.
04 Tribe Odontolochini Stebnicka & Howden, 1996
05 Tribe Odochilini Rakovic, 1987
06 Tribe Psammodiini Mulsant, 1842
07 Tribe Rhyparini Schmidt, 1910
08 Tribe Stereomerini Howden & Storey, 1992
09 Tribe Termitoderini Tangelder & Krikken, 1982
10 Tribe Thinorycterini Semenov-Tian-Shanskij & Reichardt, 1925
11 Tribe Phycochini Landin, 1960